DISCOVERY OF THE MIDDLE TRIASSIC CYCAD Nilssonia splendens IN BENXI CITY, LIAONING PROVINCE
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摘要:
尼尔桑属(Nilssonia Brongniart 1825)是苏铁植物门(Cycadophyta)代表分子之一, 地质历史时期在中国和世界均有着相当广泛的分布. 自1993年孙革报道华丽尼尔桑(Nilssonia splendens Sun 1993)以来, 该种仅见于东亚地区上三叠统. 最近, 在中国东北辽宁本溪中三叠统林家组中, 发现了一对正负面华丽尼尔桑标本, 是该种在中三叠统的首次发现. 该种的主要特征是羽叶的叶轴较细; 裂片的叶脉细而清晰, 略密, 近平行, 基本不分叉, 至顶端微上弯, 每厘米具脉约28条. 本研究根据现有标本及中国吉林天桥岭模式标本所显示的裂片和羽轴上下过渡关系, 对华丽尼尔桑的羽叶进行了重建, 并在此基础上探讨华丽尼尔桑的古地理起源与扩散. 该种在辽宁本溪林家组的发现不仅扩展了华丽尼尔桑的地质历史分布, 丰富了林家植物群面貌, 也有助于进一步了解二叠纪-三叠纪之交生物大灭绝后东北地区植物群的演替和发展.
Abstract:Nilssonia Brongniart 1825 is one of representatives of Cycadophyta. It was widely distributed in China and the world through geological time. Recently, a pair of specimens of Nilssonia splendens Sun 1993 was discovered from the Middle Triassic Linjia Formation of Benxi, Liaoning in Northeast China. The species differentiates form other species of Nilssonia in their leaves with a slender rachis; veins generally without bifurcation, clear and thinner, slightly dense, closely to parallel, slightly upward curving to the apex, occurring at concentration of 28 per cm. Since N. splendens was first identified by Sun in 1993, the species has only been documented in the Upper Triassic in East Asia. The present specimens are the first record from the Middle Triassic. According to the present fossil specimens, and the transitional relationships of laminar segments and rachis from type specimens of Tianqiaoling in Jilin, China, the reconstruction of N. splendens was suggested. The origin and diffusion of N. splendens suggest that this species from the Middle Triassic Linjia Formation, not only enlarges the knowledge in taxonomy of Nilssonia, but also provides an opportunity to understand the evolution of Nilssonia in the mass extinction through the Permian-Triassic transition and its subsequent recovery in Northeast China.
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Key words:
- Nilssonia splendens /
- Cycads /
- Middle Triassic /
- Linjia flora /
- paleogeography /
- Liaoning Province
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