Discovery, taphonomic environments and classification of Zhucheng dinosaur fauna in Shandong Province
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摘要:
诸城恐龙骨骼化石主要集中埋藏于库沟、龙骨涧、臧家庄、西见屯等化石点, 发现了鸭嘴龙、角龙、暴龙、甲龙、蜥脚类和窃蛋龙类恐龙。为进一步系统梳理诸城地区恐龙化石赋存埋藏特征, 结合前人研究和笔者近年研究发现, 诸城可能存在3种鸭嘴龙, 至少存在2种暴龙科恐龙。诸城恐龙化石最主要的埋藏相类型为泥石流沉积相。白垩纪晚期, 诸城地区干旱的气候可能是该地区恐龙死亡的原因。诸城恐龙动物群与北美地区晚白垩世恐龙组合相似, 对白垩纪恐龙的迁徙、分布、演化、生存环境、板块构造等研究提供了丰富的材料和证据。
Abstract:Kugou, Longgujian, Zangjiazhuang and Xijiantun are mainly fossil sites, and hadrosaurs, centrosaurinae, tyrannosaurs, ankylosauria, sauropods, oviraptorosauria were discovered in Zhucheng.Previous research and the author's recent studies show that there may be three kinds of hadrosaurs and at least two kinds of tyrannosauridae in Zhucheng.The main taphonomic facies type of dinosaur fossils is mud flow deposits.Draught climate may have played important roles in the mortality of dinosaurs.Zhucheng dinosaur fauna is similar to the dinosaur assemblage in North America, providing abundant materials and evidence for the study of Cretaceous dinosaurs' migration, distribution, evolution, living environment and plate tectonics.
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Key words:
- Zhucheng /
- Late Cretaceous /
- dinosaurs /
- taphonomic environment /
- Shandong Province
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图 3 山东诸城红土崖组恐龙化石埋藏层位及其岩相、沉积相特征(据旷红伟等,2014修改)
Figure 3.
表 1 诸城3种鸭嘴龙参数对比
Table 1. Parameter comparison of the three hadrosaurus in Zhucheng area
恐龙名称 发现地 荐椎数 层位 巨型山东龙 龙骨涧 10 红土崖组(比臧家庄早一点) 巨大诸城龙 龙骨涧 9 红土崖组(比臧家庄早一点) 巨大华夏龙 臧家庄 10 红土崖组 -
[1] Brinkman D B. Paleooecology of the Judith River Formation(Campanian) of Dinosaur Provincial Park, Alberta, Canada: evidence from vertebrate microfossil localities[J]. Palaeogeography, Palaeoclimatology, Palaeoecology, 1990, 78(1/2): 37-54.
[2] Brinkman D B, Russell A P, Eberth D A, et al. Vertebratepalaeo communities of the lower Judith River Group(Campanian) of south eastern Alberta, Canada, as interpreted from vertebrate microfossil assemblages[J]. Palaeogeography, Palaeoclimatology, Palaeoecology, 2004, 213: 295-313. doi: 10.1016/S0031-0182(04)00386-4
[3] Dodson P, Forster C A, Sampson S D. Ceratopsidae[C]//Weishampel D B, Dodson P, Osmólska H. The Dinosauria. Berkeley: University of California Press, 2004: 494-516.
[4] Eberth D A. Stratigraphy and sedimentology of vertebrate microfossil sites in the uppermost Judith River Formation(Campanian), Dinosaur Provincial Park, Alberta, Canada[J]. Palaeogeography Palaeoclimatology Palaeoecology, 1990, 78(1/2): 1-36.
[5] Hone D W E, Wang K B, Sullivan C, et al, A new, largetyrannosaurine theropod from the Upper Cretaceous of China[J]. Cretaceous Research, 2011, 32(4): 495-503. doi: 10.1016/j.cretres.2011.03.005
[6] He Y M, Makovicky P J, Wang K B, et al, A new leptoceratopsid(Ornithischia, Ceratopsia) with a unique ischium from the Upper Cretaceous of Shandong Province, China[J]. PLoS One, 2015a, 10(12): e0144148. doi: 10.1371/journal.pone.0144148
[7] He B Z, Qiao X F, Zhang Y L, et al., Soft-sediment deformation structures in the Cretaceous Zhucheng depression, Shandong Province, East China; their character, deformation timing and tectonic implications[J]. Journal of Asian Earth Sciences, 2015b, 110: 101-122. doi: 10.1016/j.jseaes.2014.12.005
[8] Makovicky P J. Taxonomic Revision and Phylogenetic Relationships of Basal Neoceratopsia(Dinosauria: Ornithischia)[M]. New York: Columbia University, 2002: 1-297.
[9] Sankey J T. Faunal composition and significance of high-diversity, mixedbonebeds containing Agujaceratops mariscalensis and other dinosaurs, Aguja Formation(Upper Cretaceous), Big Bend, Texas[M]. Indiana Univwesity Press, 2010: 1-47.
[10] Wood J M, Thomas R G, Visser J. Fluvial processes and vertebrate taphonomy: the upper cretaceous Judith River formation, south-central dinosaur Provincial Park, Alberta, Canada[J]. Palaeogeography, Palaeoclimatology, Palaeoecology, 1988, 66(1/2): 127-143.
[11] Xing H, Zhao X J, Wang K B, et al, Comparative osteology and phylogenetic relationship of Edmontosaurus and Shantungosaurus(Dinosauria: Hadrosauridae) from the Upper Cretaceous of North America and East Asia[J]. Acta Geologica Sinica, 2014, 88(6): 1801-1840.
[12] Xu X, Wang K B, Zhao X J, et al. First ceratopsid dinosaur from China and its biogeographical implications[J]. Chinese Science Bulletin, 2010a, 55(16): 1631-1635. doi: 10.1007/s11434-009-3614-5
[13] Xu X, Wang K B, Zhao X J, et al, A newleptoceratopsid(Ornithischia: Ceratopsia) from the Upper Cretaceous of Shandong, China and its implications for neoceratopsian evolution[J]. PLoS One, 2010b, 5(11): e13835. doi: 10.1371/journal.pone.0013835
[14] Yu Y L, Wang K B, Chen S Q, et al, A new caenagnathid dinosaur from the Upper Cretaceous Wangshi Group of Shandong, China, with comments on size variation among oviraptorosaurs[J]. Scientific Reports, 2018, 8(1): 1-10.
[15] 安伟, 旷红伟, 柳永清, 等. 山东诸城晚白垩世王氏群恐龙化石层碎屑锆石定年和物源示踪[J]. 地质论评, 2016, 62(2): 453-471. doi: 10.16509/j.georeview.2016.02.017
[16] 何碧竹, 乔秀夫, 田洪水, 等. 山东诸城晚白垩世古地震事件与恐龙化石埋藏[J]. 古地理学报, 2011, 13(6): 615-626. https://www.cnki.com.cn/Article/CJFDTOTAL-GDLX201106007.htm
[17] 胡承志. 山东诸城巨型鸭嘴龙化石[J]. 地质学报, 1973, 2: 179-206. doi: 10.19762/j.cnki.dizhixuebao.1973.02.002
[18] 胡承志, 程政武. 巨型山东龙再研究的新进展[J]. 地球学报, 1986, 14: 163-170. https://www.cnki.com.cn/Article/CJFDTOTAL-DQXB198603011.htm
[19] 胡承志, 程政武, 庞其清, 等. 巨型山东龙[M]. 北京: 地质出版社, 2001: 1-139.
[20] 季燕南. 巨型山东龙的系统分类, 生活习性与生态环境研究[J]. 地学前缘, 2010a, (1): 378-385. https://www.cnki.com.cn/Article/CJFDTOTAL-DXQY201001038.htm
[21] 季燕南. 山东省诸城地区晚白垩世沉积环境与山东龙集群埋藏研究[D]. 中国地质大学(北京) 硕士学位论文, 2010b: 1-2.
[22] 旷红伟, 许克民, 柳永清, 等. 胶东诸城晚白垩世恐龙骨骼化石地球化学及埋藏学研究[J]. 地质论评, 2013, 59(6): 1001-1023. https://www.cnki.com.cn/Article/CJFDTOTAL-DZLP201306001.htm
[23] 旷红伟, 柳永清, 董超, 等. 山东诸城晚白垩世恐龙化石埋藏学研究[J]. 地质学报, 2014, 88(8): 1353-1371. https://www.cnki.com.cn/Article/CJFDTOTAL-DZXE201408001.htm
[24] 卢磊勋, 杜萌, 邵瀚石. K-Ar法和Ar-Ar法两种定年方法的差异性对比及讨论[J]. 辽宁化工, 2015, (8): 1025-1027. https://www.cnki.com.cn/Article/CJFDTOTAL-LNHG201508042.htm
[25] 柳永清, 旷红伟, 姬书安, 等. 晚中生代地球表层重大地质事件的陆地环境剧变与生物群演替响应[J]. 地学前缘, 2009, 16(5): 37-47. https://www.cnki.com.cn/Article/CJFDTOTAL-DXQY200905005.htm
[26] 柳永清, 旷红伟, 彭楠, 等. 鲁东诸城地区晚白垩世恐龙集群埋藏地沉积相与埋藏学初步研究[J]. 地质论评, 2010, 56(4): 457-467. https://www.cnki.com.cn/Article/CJFDTOTAL-DZLP201004000.htm
[27] 柳永清, 旷红伟, 彭楠, 等. 山东胶莱盆地白垩纪恐龙足迹与骨骼化石埋藏沉积相与古地理环境[J]. 地学前缘, 2011, 18(4): 9-24. https://www.cnki.com.cn/Article/CJFDTOTAL-DXQY201104001.htm
[28] 毛兴强, 王恩德, 杨群, 等. 山东省胶东半岛新立金矿床成因[J]. 地质通报, 2022, 41(10): 1855-1868. http://dzhtb.cgs.cn/gbc/ch/reader/view_abstract.aspx?file_no=20221013&flag=1
[29] 孟繁聪, 李天福, 薛怀民, 等. 胶莱盆地晚白垩世不同地幔源区的两种基性岩浆——诸城玄武岩和胶州玄武岩的对比[J]. 岩石学报, 2006, 22(6): 1644-1656. https://www.cnki.com.cn/Article/CJFDTOTAL-YSXB200606021.htm
[30] 莫进尤, 王克柏, 陈树清, 等. 山东晚白垩世一新的巨龙类恐龙[J]. 地质通报, 2017, 36(9): 1501-1505. http://dzhtb.cgs.cn/gbc/ch/reader/view_abstract.aspx?file_no=20170901&flag=1
[31] 王能盛, 旷红伟, 柳永清, 等. 中国东部晚白垩世恐龙化石集群埋藏特征及国内外对比[J]. 古地理学报, 2015, 17(5): 593-610. https://www.cnki.com.cn/Article/CJFDTOTAL-GDLX201505002.htm
[32] 王克柏, 张艳霞, 陈军, 等. 山东诸城地区晚白垩世一新的甲龙类恐龙[J]. 地质通报, 2020, 39(7): 958-962. http://dzhtb.cgs.cn/gbc/ch/reader/view_abstract.aspx?file_no=20200703&flag=1
[33] 闫峻, 陈江峰, 谢智, 等. 鲁东晚白垩世玄武岩中的幔源捕虏体: 对中国东部岩石圈减薄时间制约的新证据[J]. 科学通报, 2003, 48(14): 1570-1574. https://www.cnki.com.cn/Article/CJFDTOTAL-KXTB200314017.htm
[34] 闫峻, 陈江峰, 谢智, 等. 鲁东白垩世玄武岩及其中幔源包体的岩石学和地球化学研究[J]. 岩石学报, 2005, 21(1): 99-112. https://www.cnki.com.cn/Article/CJFDTOTAL-YSXB200501011.htm
[35] 赵喜进, 李敦景, 韩岗, 等. 山东的巨大诸城龙[J]. 地球学报, 2007, 28(2): 111-122. https://www.cnki.com.cn/Article/CJFDTOTAL-DQXB200702002.htm
[36] 赵喜进, 李敦景. 巨大诸城龙[M]. 北京: 北京艺术与科学电子出版社, 2008: 1-222.
[37] 赵喜进, 王克柏, 李敦景. 巨大华夏龙[J]. 地质通报, 2011, 31(11): 1671-1688. http://dzhtb.cgs.cn/gbc/ch/reader/view_abstract.aspx?file_no=20111103&flag=1
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